Nitrogen signaling factor triggers a respiration-like gene expression program in fission yeast

EMBO J. 2024 Oct;43(20):4604-4624. doi: 10.1038/s44318-024-00224-z. Epub 2024 Sep 10.

Abstract

Microbes have evolved intricate communication systems that enable individual cells of a population to send and receive signals in response to changes in their immediate environment. In the fission yeast Schizosaccharomyces pombe, the oxylipin nitrogen signaling factor (NSF) is part of such communication system, which functions to regulate the usage of different nitrogen sources. Yet, the pathways and mechanisms by which NSF acts are poorly understood. Here, we show that NSF physically interacts with the mitochondrial sulfide:quinone oxidoreductase Hmt2 and that it prompts a change from a fermentation- to a respiration-like gene expression program without any change in the carbon source. Our results suggest that NSF activity is not restricted to nitrogen metabolism alone and that it could function as a rheostat to prepare a population of S. pombe cells for an imminent shortage of their preferred nutrients.

Keywords: Cell-to-Cell Communication; Fission Yeast; Mitochondrial Respiration; Nitrogen Catabolite Repression; Nitrogen Signaling Factor.

MeSH terms

  • Gene Expression Regulation, Fungal*
  • Nitrogen* / metabolism
  • Schizosaccharomyces pombe Proteins* / genetics
  • Schizosaccharomyces pombe Proteins* / metabolism
  • Schizosaccharomyces* / genetics
  • Schizosaccharomyces* / metabolism
  • Signal Transduction*

Substances

  • Schizosaccharomyces pombe Proteins
  • Nitrogen