Human leukocyte antigen class I antibody-activated endothelium promotes CD206+ M2 macrophage polarization and MMP9 secretion through TLR4 signaling and P-selectin in a model of antibody-mediated rejection and allograft vasculopathy

Am J Transplant. 2024 Mar;24(3):406-418. doi: 10.1016/j.ajt.2023.10.020. Epub 2023 Oct 29.

Abstract

HLA donor-specific antibodies (DSA) elicit alloimmune responses against the graft vasculature, leading to endothelial cell (EC) activation and monocyte infiltration during antibody-mediated rejection (AMR). AMR promotes chronic inflammation and remodeling, leading to thickening of the arterial intima termed transplant vasculopathy or cardiac allograft vasculopathy (CAV) in heart transplants. Intragraft-recipient macrophages serve as a diagnostic marker in AMR; however, their polarization and function remain unclear. In this study, we utilized an in vitro Transwell coculture system to explore the mechanisms of monocyte-to-macrophage polarization induced by HLA I DSA-activated ECs. Anti-HLA I (IgG or F(ab')2) antibody-activated ECs induced the polarization of M2 macrophages with increased CD206 expression and MMP9 secretion. However, inhibition of TLR4 signaling or PSGL-1-P-selectin interactions significantly decreased both CD206 and MMP9. Monocyte adherence to Fc-P-selectin coated plates induced M2 macrophages with increased CD206 and MMP9. Moreover, Fc-receptor and IgG interactions synergistically enhanced active-MMP9 in conjunction with P-selectin. Transcriptomic analysis of arteries from DSA+CAV+ rejected cardiac allografts and multiplex-immunofluorescent staining illustrated the expression of CD68+CD206+CD163+MMP9+ M2 macrophages within the neointima of CAV-affected lesions. These findings reveal a novel mechanism linking HLA I antibody-activated endothelium to the generation of M2 macrophages which secrete vascular remodeling proteins contributing to AMR and CAV pathogenesis.

Keywords: antibody-mediated rejection (AMR); areas of interest (AOIs); cardiac allograft vasculopathy (CAV); donor-specific HLA antibodies (DSA); endothelial cell (EC); human leukocyte antigen (HLA); transplant vasculopathy (TV).

MeSH terms

  • Allografts
  • Endothelium
  • HLA Antigens
  • Humans
  • Immunoglobulin G
  • Macrophages
  • Matrix Metalloproteinase 9
  • P-Selectin
  • Toll-Like Receptor 4*
  • Vascular Diseases*

Substances

  • Toll-Like Receptor 4
  • Matrix Metalloproteinase 9
  • P-Selectin
  • HLA Antigens
  • Immunoglobulin G
  • TLR4 protein, human
  • MMP9 protein, human