Dancr-BRG1 regulates Nfatc1 transcription and Pgc1β-dependent metabolic shifts in osteoclastogenesis

Proc Natl Acad Sci U S A. 2024 Jan 30;121(5):e2313656121. doi: 10.1073/pnas.2313656121. Epub 2024 Jan 22.

Abstract

Long non-coding RNA (lncRNA) serves as a vital regulator of bone metabolism, but its role in pathologically overactive osteoclast differentiation remains elusive. Here, we identify lncRNA Dancr (Differentiation Antagonizing Non-protein Coding RNA) as a critical suppressor of osteoclastogenesis and bone resorption, which is down-regulated in response to estrogen deficiency. Global or osteoclast-specific Dancr Knockout mice display significant trabecular bone deterioration and enhanced osteoclast activity, but minimal alteration of bone formation. Moreover, the bone-targeted delivery of Dancr by Adeno-associated viral remarkably attenuates ovariectomy-induced osteopenia in mice. Mechanistically, Dancr establishes a direct interaction with Brahma-related gene 1 to prevent its binding and preserve H3K27me3 enrichment at the nuclear factor of activated T cells 1 and proliferator-activated receptor gamma coactivator 1-beta promoters, thereby maintaining appropriate expression of osteoclastic genes and metabolic programs during osteoclastogenesis. These results demonstrate that Dancr is a key molecule maintaining proper osteoclast differentiation and bone homeostasis under physiological conditions, and Dancr overexpression constitutes a potential strategy for treating osteoporosis.

Keywords: Dancr; LncRNA; bone resorption; osteoclast; osteoporosis.

MeSH terms

  • Animals
  • Female
  • Homeostasis
  • Mice
  • Mice, Knockout
  • NFATC Transcription Factors* / genetics
  • Osteoclasts
  • Osteogenesis* / genetics
  • RNA, Long Noncoding* / genetics
  • Transcription Factors* / genetics

Substances

  • NFATC Transcription Factors
  • Nfatc1 protein, mouse
  • RNA, Long Noncoding
  • Transcription Factors
  • Smarca4 protein, mouse