1 1NIOZ Royal Netherlands Institute for Sea Research, Department of Coastal Systems, and Utrecht University, P.O. Box 59, 1790 AB Den Burg, Texel The Netherlands.
2 2Centre for Biological Diversity, School of Biology, University of St Andrews, Fife, KY16 9TF UK.
3 3Rudi Drent Chair in Global Flyway Ecology, Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences (GELIFES), University of Groningen, PO Box 11103, 9700 CC Groningen, The Netherlands.
4 4EBIOME Ecobiologie Marine et Environnement, Département de Biologie, L'université de Nouakchott Al-Aasriya, BP 880 Nouakchott, Mauritania.
5 5Present Address: Sorbonne Université, CNRS, Station Biologique de Roscoff, Laboratoire Adaptation et Diversité en Milieu Marin, UMR 7144, CS90074, 29688 Roscoff Cedex, France.