Rap1 promotes endothelial mechanosensing complex formation, NO release and normal endothelial function

EMBO Rep. 2015 May;16(5):628-37. doi: 10.15252/embr.201439846. Epub 2015 Mar 25.

Abstract

Decreased nitric oxide (NO) bioavailability underlies a number of cardiovascular pathologies, including hypertension. The shear stress exerted by flowing blood is the main determinant of NO release. Rap1 promotes integrin- and cadherin-mediated signaling. Here, we show that Rap1 is a critical regulator of NO production and endothelial function. Rap1 deficiency in murine endothelium attenuates NO production and diminishes NO-dependent vasodilation, leading to endothelial dysfunction and hypertension, without deleterious effects on vessel integrity. Mechanistically, Rap1 is activated by shear stress, promotes the formation of the endothelial mechanosensing complex-comprised of PECAM-1, VE-cadherin and VEGFR2- and downstream signaling to NO production. Our study establishes a novel paradigm for Rap1 as a regulator of mechanotransduction.

Keywords: mechanotransduction; nitric oxide; shear stress; small GTPase Rap1; vasodilation.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Blood Pressure
  • Capillary Permeability / genetics
  • Endothelium / metabolism*
  • Humans
  • Hypertension / genetics
  • Hypertension / metabolism
  • Hypertension / physiopathology
  • Hypertrophy, Left Ventricular / genetics
  • Hypertrophy, Left Ventricular / metabolism
  • Hypertrophy, Left Ventricular / physiopathology
  • Male
  • Mechanotransduction, Cellular*
  • Mice
  • Mice, Knockout
  • Models, Biological
  • Nitric Oxide / metabolism*
  • Nitric Oxide Synthase Type III / metabolism
  • Organ Specificity / genetics
  • Signal Transduction
  • Vasodilation / genetics
  • rap1 GTP-Binding Proteins / genetics
  • rap1 GTP-Binding Proteins / metabolism*

Substances

  • Nitric Oxide
  • Nitric Oxide Synthase Type III
  • rap1 GTP-Binding Proteins