1 CSIRO Ecosystem Sciences, , Canberra, Australian Capital Territory, Australia, Department of Life Sciences, Natural History Museum, , Cromwell Road, London SW7 5BD, UK, UC Davis Genome Center, , Davis, CA 95616, USA, Department of Ecology and Evolutionary Biology, University of Arizona, , Tucson, AZ 85721, USA, Institute of Evolutionary Biology, University of Edinburgh, , Edinburgh EH9 3JT, UK, Research School of Biology, Australian National University, , Canberra, Australian Capital Territory 0200, Australia, Department of Biology, Ghent University, , Marine Biology Lab, Ghent 9000, Belgium, Department of Biology, New York University, , New York, NY 10003, USA, National Research Council, Institute of Ecosystem Study, , Largo Tonolli 50, 28922 Verbania Pallanza, Italy, Department of Genetics and Evolution, University of Geneva, , Sciences III, 30, Quai Ernest Ansermet, Geneva 1211, Switzerland, Department of Chemistry and Biochemistry and BioFrontiers Institute, University of Colorado, , Boulder, CO 80309, USA, Department of Life Sciences, Imperial College London, , Ascot, Berkshire SL5 7PY, UK, Hubbard Center for Genome Studies, University of New Hampshire, , 35 Colovos Road, Durham, NH 03824, USA, School of Biological Sciences, Bangor University, , Environment Centre Wales Building, Deiniol Road, College of Natural Sciences, Gwynedd LL57 2UW, UK.