Conservation of the metabolomic response to starvation across two divergent microbes

Proc Natl Acad Sci U S A. 2006 Dec 19;103(51):19302-7. doi: 10.1073/pnas.0609508103. Epub 2006 Dec 11.

Abstract

We followed 68 cellular metabolites after carbon or nitrogen starvation of Escherichia coli and Saccharomyces cerevisiae, using a filter-culture methodology that allows exponential growth, nondisruptive nutrient removal, and fast quenching of metabolism. Dynamic concentration changes were measured by liquid chromatography-tandem mass spectrometry and viewed in clustered heat-map format. The major metabolic responses anticipated from metabolite-specific experiments in the literature were observed as well as a number of novel responses. When the data were analyzed by singular value decomposition, two dominant characteristic vectors were found, one corresponding to a generic starvation response and another to a nutrient-specific starvation response that is similar in both organisms. Together these captured a remarkable 72% of the metabolite concentration changes in the full data set. The responses described by the generic starvation response vector (42%) included, for example, depletion of most biosynthetic intermediates. The nutrient-specific vector (30%) included key responses such as increased phosphoenolpyruvate signaling glucose deprivation and increased alpha-ketoglutarate signaling ammonia deprivation. Metabolic similarity across organisms extends from the covalent reaction network of metabolism to include many elements of metabolome response to nutrient deprivation as well.

Publication types

  • Comparative Study
  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't
  • Research Support, U.S. Gov't, Non-P.H.S.

MeSH terms

  • Carbon / deficiency*
  • Chromatography, Liquid
  • Cluster Analysis
  • Energy Metabolism / physiology*
  • Escherichia coli K12 / metabolism
  • Escherichia coli K12 / physiology*
  • Escherichia coli Proteins / metabolism*
  • Mass Spectrometry
  • Nitrogen / deficiency*
  • Phenylpyruvic Acids / metabolism
  • Saccharomyces cerevisiae / metabolism
  • Saccharomyces cerevisiae / physiology*
  • Saccharomyces cerevisiae Proteins / metabolism*
  • Species Specificity

Substances

  • Escherichia coli Proteins
  • Phenylpyruvic Acids
  • Saccharomyces cerevisiae Proteins
  • Carbon
  • Nitrogen
  • phenylpyruvic acid